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The first appearance of trilobites in the fossil record defines the base of the Atdabanian stage of the Early Cambrian period (), and they flourished throughout the lower Paleozoic era before beginning a drawn-out decline to extinction when, during the Devonian, all trilobite orders except the Proetids died out.Trilobites disappeared in the mass extinction at the end of the Permian about 252 million years ago.Most lifestyles expected of modern marine arthropods are seen in trilobites, with the possible exception of parasitism (where scientific debate continues).All Olenellina lack facial sutures (see below), and this is thought to represent the original state.Silurian and Devonian trilobite assemblages are superficially similar to Ordovician assemblages, dominated by Lichida and Phacopida (including the well-known Calymenina).

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include the origin of new types of eyes, improvement of enrollment and articulation mechanisms, increased size of pygidium (micropygy to isopygy), and development of extreme spinosity in certain groups.

meaning "three lobes") are a group of extinct marine arachnomorph arthropods that form the class Trilobita.



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